Large amounts of chondroitin sulfate turn cartilage into a hard gel. When the tissue shrinks during dehydration it does so by enlarging the holes (lacunae) around the cells. Tendons have much less ground substance gluing the bundles of fibers together. When they shrink the bundles of fibers can simply separate from one another. Since the fibroblasts lie between the bundles of collagen instead of within them the nuclei are seen in cracks between the fibers.